2025
Bapat, A., Jain, V., Blum, C.R., Boucherie, P.H., Sumasgutner, P., Bugnyar,T., 2026. Spatio-temporal patterns of juvenile common ravens integrating into a free-flying non-breeder flock,iScience (, doi: https://doi.org/10.1016/j.isci.2025.114412
Šlipogor, V., Masilkova M., Höflinger A., Lang N., Bugnyar T., Konečná M., 2025. With or without you: common marmoset, Callithrix jacchus, personality expression is mediated by social setting, Animal Behaviour, https://doi.org/10.1016/j.anbehav.2025.123367.
Weißenborn, J.A.E., Sumasgutner, P., Šlipogor, V. and Bugnyar, T. 2025. Startling ravens Corvus corax at foraging: differences in anti-predator behaviour can be explained by age rather than personality. J Avian Biol, 2025: e03437. https://doi.org/10.1002/jav.03437
Boucherie, P.H., Murari, G., Svitil, E.A., Bugnyar, T. 2025. Rearing group size in young ravens: stress load or social opportunity? R. Soc.Open Sci. 12: 250479
Damini, S., Blum, C.R., Sumasgutner, P. et al. 2025. When to mob? plasticity of antipredator behavior in common ravens’ families (Corvus corax) across offspring development. Anim Cogn 28, 55. https://doi.org/10.1007/s10071-025-01976-9
Bapat, A., Kempf, A., Friry, S., Boucherie P.H., Bugnyar, T. 2025. Patters of object play behaviour and its functional implication in free-flying common ravens. Scientific Reports. https://doi.org/15.10.1038/s41598-024-83856-9.
Vanhooland, L.-C. & Mager, C. & Teuben, A. & Bugnyar, T. & Massen, J.M.M. 2025. Comparing the performances of chimpanzees (Pan troglodytes) and gorillas (Gorilla gorilla gorilla) in two self-awareness tasks. American Journal of Primatology. 87. 10.1002/ajp.70010.
Bugnyar, T. & Klump, B. 2025. Avian social learning. In: Encyclopedia of Animal Behaviour, Third Edition, Springer.
Ye, H., Šlipogor, V., & Hanson, B., Séneca, J.,Hausmann, B., Herbold, C., Pjevac, P., Bugnyar, T., Loy, A. 2025. Associations between gut microbiota and personality traits: insights from a captive common marmoset (Callithrix jacchus) colony. 10.1101/2025.02.12.637913.
Miller, R., Boeckle, M., Ridgway, S., Richardson, J., Uhl, F., Bugnyar, T., Schwab, C. 2025. Social attention across development in common ravens and carrion/hooded crows. Animal Behaviour. 220. 123038. 10.1016/j.anbehav.2024.11.016.
Massen, J.M.M., Boucherie P.H., Bugnyar, T. 2025. Third-party interventions of common ravens, Corvus corax, Animal Behaviour, https://doi.org/10.1016/j.anbehav.2024.10.018
2024
Bugnyar, T. 2024. Why are ravens smart? Exploring the social intelligence hypothesis. Journal of Ornithology. https://doi.org/10.1007/s10336-023-02111-6
Horn, L., Bugnyar, T. & Massen, J.J.M. 2024. Testing the effects of kinship, reciprocity and dominance on prosocial food provisioning in azure-winged magpies (Cyanopica cyana), carrion crows (Corvus corone) and common ravens (C. corax). Ethology. 130, 4, e13420. https://doi.org/10.1111/eth.1342
Horn, L. Windhager, S., Juricka, N., Bugnyar, T., Massen, J.J.M. & Markova, G. 2024. Investigating the effects of prenatal testosterone exposure (via 2D:4D) and socio-relational factors on 3-6-year-old preeschoolers’ prosocial choices. Early Human Development, 195, 106055. https://doi.org/10.1016/j.earlhumdev.2024.106055
Suwandschieff, E., Huber, L., Bugnyar, T. & Schwing, R., 2024. Kea, bird of versatility. Kea parrot (Nestor notabilis) show high behavioral flexibility in solving a demonstrates sequence task. Journal of Ornithology. https://doi.org/10.1007/s10336-023-02127-y
2023
Suwandschieff, E. & Wein, A. & Folkertsma, R. & Bugnyar, T. & Huber, L. & Schwing, R. 2023. Two-action task, testing imitative social learning in kea (Nestor notabilis). Animal Cognition. 26, 4, S. 1395-1408 14 S. doi: 10.1007/s10071-023-01788-9.
Federspiel, I. & Schmitt, V. & Schuster, R. & Rockenbach, C. & Braun, A. & Loretto, M.C. & Michels, C. & Fischer, J. & Mussweiler, T. & Bugnyar, T. 2023. Are You Better Than Me? Social Comparisons in Carrion Crows (Corvus corone). https://doi.org/10.21203/rs.3.rs-2500378/v1
Vanhooland, L-C., Szabó, A., Bugnyar, T. & Massen, J. J. M., 2023. A comparative study of mirror self-recognition in three corvid species. Animal Cognition. 26, 1, S. 229-248 20 S. https://doi.org/10.1007/s10071-022-01696-4
2022
Aguenounoun, G., Bugnyar, T, & Keupp, S. 2022. The evolution of primate short term memory. Animal Behaviour and Cognition, 9 (4). 10.26451/abc.09.04.06.2022
Blum, C.R., Fitch, W.T.S., & Bugnyar, T. 2022. Social dynamics impact scolding behaviour in captive groups of common ravens (Corvus corax). Frontiers in Zoology, 19 (1), 1-12.
Boucherie, P.H., Gallego-Abenza, M., Massen, J.J.M. & Bugnyar, T. 2022. Dominance in a socially dynamic setting: hierarchical structure and conflict dynamics in raven foraging groups. Philosophical Transactions of the Royal Society Series B, 377, 20200446. https://doi.org/10.1098/rstb.2020.0446
Gallego-Abenza, M., Boucherie, P.H. & Bugnyar, T. 2022: Early social environment affects the attention to social cues in juvenile common ravens, Corvus corax. Royal Society Open Science, 9 (6), 220132.
Gallup, A.C., Schild, A.B., Ühlein, M.A., Bugnyar, T. & Massen, J.J.M. 2022. No evidence for contagious yawning in juvenile ravens (Corvus corax): An observational study. Animals, 12 (11), 1357.
Horn, L., Cimarelli, G., Boucherie, P.H., Slipogor, V. & Bugnyar, T. 2022. Beyond the dichotomy between field and lab – the importance of studying cognition in context. Current opinion in Behavioral Sciences, 46, 101172.
Jain, V., Bugnyar, T., Cunningham, S.J., Gallego-Abenza, M., Loretto, M-C. & Sumasgutner, P. 2022. The spatial and temporal exploitation of anthrpogenic food sources by common ravens (Corvus corax) in the Alps. 2022. Movement Ecology, 10, 35. https://doi.org/10.1186/s40462-022-00335-4
Masilkova, M., Slipogor, V., Lima Marques Silva, G.H., Hadova, M., Lhota, S., Bugnyar, T. & Konecna, M. 2022. Age, but not hand preference, is related to personality traits in common marmosets (Callithrix jacchus). Royal Society Open Science, 9, 220797.
Miller, R., Lambert, M.L., Frohnwieser, A., Brecht, K.F., Bugnyar, T., Crampton, I., Garcia-Pelegrin, E., Gould, K., Greggor, A.L., Izawa, E.-I., Kelly, D.M., Li, Z., Luo, Y., Luong, L.B., Massen, J.J.M., Nieder, A., Reber, S.A., Schiestl, M., Seguchi, A., Sepehri, P., Stevens, J.R., Taylor, A.H., Wang, L., Wolff, L.M., Zhang, Y. & Clayton, N.S. 2022. Socio-ecological correlates of neophobia in corvids. Current Biology, 32 (1), 74-85.
Slipogor, V., Graf, C., Massen, J.J.M., & Bugnyar, T. 2022. Personality and social context predict cognitive performance in common marmosets (Callithrix jacchus). Scientific Reports, 12, 6702.
Vanhooland, L.-C., Szabo, A., Bugnyar, T. & Massen, J.J.M. 2022. A comparative study on mirror self-recognition in three corvid species. Animal Cognition, https://doi.org/10.1007/s10071-022-01696-4
Wenig, K., Pacher, L. & Bugnyar, T. 2022. Testing the contagious nature of allopreening: bystander ravens are affected by conspecifics’ affiliative interactions. Animal Behaviour, 184, 71-80.
2021
Adriaense, J.E.C., Šlipogor, V., Hintze, S. Marshall, L., Lamm, C. & Bugnyar, T. 2021. Watching others in a positive state does not induce optimism bias in common marmosets (Callithrix jacchus), but leads to behaviour indicative of competition. Animal Cognition, 24, 1039-1056.
Bugnyar, T. 2021. Raven social cognition and behavior. In: Kaufman, A.B., Call, J., Kaufman, J.C. (eds.) The Cambridge Handbook of Animal Cognition, pp. 322-342, Cambridge, Cambridge University Press. https://doi.org/10.1017/9781108564113.018
Ersoy, S., Maag, N., Boehly, T., Boucherie, P.H. & Bugnyar, T. 2021. Sex-specific parental care during post-fledging in common ravens. Animal Behaviour, 181, 95-103. https://doi.org/10.1016/j.anbehav.2021.09.004
Gallego-Abenza, M., Blum, C. & Bugnyar, T. 2021. Who is crying wolf? Seasonal effect on anti-predator response to age-specific alarm-calls in common ravens, Corvus corax. Learning & Behavior, https://doi.org/10.3758/s13420-020-00455-0
Horn, L., Zewald, J.S., Bugnyar, T & Massen, J.J.M. 2021. Carrion crows and azure-winged magpies show no prosocial tendencies when tested in a token transfer paradigm. Animals, 11(6), 1526. https://doi.org/10.3390/ani11061526
Keupp, S., Abedin, F., Jeanson, L., Kade, C., Kalbitz, J., Titchener, R., Mussweiler, T., Bugnyar, T. & Fischer, J. 2021. Performance-based social comparisons in humans and long-tailed macaques. Animal Behavior and Cognition, 8(3), 325-350. https://doi.org/10.26451/abc.08.03.02.2021
Martin, J.S., Koski, S.E, Bugnyar, T., Jaeggi, A.V. & Massen, J.J.M. (2021). Prosociality, social tolerance and partner choice facilitate mutually beneficial cooperation in common marmosets, Callithrix jacchus. Animal Behaviour, 173, 115-136.
Massen, J.J.M., Hartlieb, M., Martin, J.S., Leitgeb, E.B., Hockl, J., Kocourek, M., Olkowicz, S., Zhang, Y., Osadnik, C., Verkleij, J.W., Bugnyar, T., Nemec, P., & Gallup A.C. 2021. Brain site and neuron numbers drive differences in yawn duration across mammals and birds. Communications Biology, 4, 503. https://doi.org/10.1038/s42003-021-02019-y | http://www.nature.com/commsbio
Miller, R., Lambert, M.L., Frohnwieser, A., Brecht, K.F., Bugnyar, T., Crampton, I., Garcia-Pelegrin, E., Gould, K., Greggor, A.L., Izawa, E.-I., Kelly, D.M., Li, Z., Luo, Z., Luong, L.B., Massen, J.J.M., Nieder, A., Reber, S.A., Schiestl, M., Seguchi, A., Sepehri, P., Stevens, J.R., Taylor, A.H., Wang, L., Wolff, L.M., Zhang, Y. & Clayton, N.S. 2021. Socio-ecological correlates of neophobia in corvids. Current Biology, published online Nov 17.
Šlipogor, V., Massen, J.J.M., Schiel, N., Souto, A. & Bugnyar, T. 2021. Temporal consistency and ecological validity of personality structure in common marmosets (Callithrix jacchus): A unifying field and laboratory approach. American Journal of Primatology, 83, e23229. https://doi.org/10.1002/ajp.23229
Stocker, M., Prosl, J., Vanhooland, L.-C., Horn, L., Bugnyar, T., Canoine, V. & Massen, J.J.M. 2021. Measuring salivary mesotocin in birds. Seasonal differences in ravens’ peripheral mesotocin levels. Hormones & Behaviour, 134, 105015. https://doi.org/10.1016/j.yhbeh.2021.105015
Wenig, K., Boucherie, P.H., & Bugnyar, T. 2021. Early evidence for emotional play contagion in juvenile ravens. Animal Cognition, 24, 717–729. https://doi.org/10.1007/s10071-020-01466-0
2020
Beck, K.B., Loretto, M.-C. & Bugnyar, T. 2020. Effects of site fidelity, group size and age on food-caching behaviour of common ravens, Corvus corax. Animal Behaviour, 164, 51-64. https://doi.org/10.1016/j.anbehav.2020.03.015
Blum, C.R., Fitch, W.T. & Bugnyar, T. 2020. Rapid learning and long-term memory for dangerous humans in ravens (Corvus corax). Frontiers in Psychology, 11, 2843.
Boucherie, P.H., Blum, C. & Bugnyar, T. 2020. Effect of rearing style on the development of social behaviour in young ravens (Corvus corax). Ethology, 126 (6), 595-609. https://doi.org/10.1111/eth.13010
Davídková, M., Veselý, P., Syrová, M., Nácarová, J. & Bugnyar, T. 2020. Ravens respond to unfamiliar corvid alarm calls. Journal of Ornithology, 161 (4), 967-975. 10.1007/s10336-020-01781-w
Gallego-Abenza, M., Loretto, M.-C. & Bugnyar, T. 2020. Decision time modulates social foraging success in wild common ravens, Corvus corax. Ethology, 126 (4), 413-422. https://doi.org/10.1111/eth.12986
Horn, L., Bugnyar, T., Griesser, M., Hengl, M., Izawa, E.-I., Oortwijn, T., Rössler, C., Scheer, C., Schiestl, M., Suyama, M., Taylor, A.H., Vanhooland, L.-C., von Bayern, A.M.P., Zürcher, Y. & Massen, J.J.M. (2020). Sex-specific effects of cooperative breeding and colonial nesting on prosociality in corvids. eLife, 9, e58139. DOI: 10.7554/eLife.58139
Kirchmeir, A., Uhl, F., Bugnyar, T., & Schwab, C. 2020. Catching crows: seasonality, techniques and the influence of social behaviour. Ringing & Migration, 1-7. https://doi.org/10.1080/03078698.2019.1759906
Loretto, M.-C., Schuster, R., Federspiel, I.G., Heinrich, B. & Bugnyar, T. 2020. Contextual imitation in juvenile common ravens (Corvus corax). Animal Behaviour, 163, 127-134. https://doi.org/10.1016/j.anbehav.2020.03.007
Massen, J.J.M., Haley, S.M., & Bugnyar, T. (2020). Azure-winged magpies‘ decisions to share are contingent on the presence or absence of food for the recipient. Scientific Reports, 10 (1), 16147. 10.1038/s41598-020-73256-0
Massen, J.J.M., Bauer, L., Spurny, B., Bugnyar, T. & Kret, M.E. (2020). Reply to: ‚The data do not support the existence of an ‘old boy network’ in science. Some critical comments on a study by Massen et al. Scientific Reports, 10, 13783. 10.1038/s41598-020-69108-6
Massen, J.J.M., Schaake, W. & Bugnyar, T. (2020). A comparison of cooperative cognition in corvids, chimpanzees, and other animals. In: Hopper, L.M. & Ross, S.R. (eds) Chimpanzees in Context. A Comparative perspective of Chimpanzee Behavior, Cognition, Conservation, and Welfare. pp 368-390; Chicago, University of Chicago Press.
Morales Picard, A., Mundry, R., Auersperg, A.M.I., Boeving, E.R., Boucherie, P., Bugnyar, T., Emery, N., Federspiel, I.G., Gajdon, G.K., Guery, J.-P., Hegedic, M., Horn, L., Kavanagh, E., Lambert, M., Massen, J.J.M., Rodrigues, M., Schiestl, M., Schwing, R., Szabo, B., Taylor, A., van Horik, J., von Bayern, A., Seed, A. & Slocombe, K. 2019. Why preen others? Predictors of allopreening in parrots and corvids and comparisons to grooming in great apes. Ethology, 126 (2), 207-228. https://doi.org/10.1111/eth.12999.
Sierro, J., Loretto, M.-C., Szipl, G., Massen, J.J.M., & Bugnyar, T. (2020). Food calling in ravens (Corvus corax) revisited: who is addressed? Ethology, 126 (2), 257-266. https://doi.org/10.1111/eth.12991
Šlipogor, V., Burkart, J.M., Martin, J.S., Bugnyar, T., & Koski, S.E. (2020). Personality method validation in common marmosets (Callithrix jacchus): Getting the best of both worlds. Journal of Comparative Psychology, 134 (1), 52-70. 10.1037/com0000188
Stocker, M., Loretto, M.-C., Sterck, E.H.M., Bugnyar, T. & Massen J.J.M. 2020. Cooperation with closely bonded individuals reduced cortisol levels in long-tailed macaques. Royal Society Open Science, 7 (5), 191056. 10.1098/rsos.191056
Vanhooland, L.-C., Bugnyar, T. & Massen, J.J.M. 2020. Crows (Corvus corone ssp.) check contingency in a mirror yet fail the mirror-mark test. Journal of Comparative Psychology, 134 (2), 158-169. 10.1037/com0000195
Wascher, C.A.F, Feider, B., Bugnyar, T. & Dufour, V. 2019. Crows and common ravens do not reciprocally exchange tokens with a conspecific to gain food rewards. Ethology, 126 (2), 278-287. https://doi.org/10.1111/eth.12970
2019
Adriaense, J.E.C., Martin, J.S., Schiestl, M., Lamm, C. & Bugnyar, T. 2019. Negative emotional contagion and cognitive bias in common ravens (Corvus corax). Proceedings of the National Academy of Sciences of the United States of America, 116, 11547-11552. 10.1073/pnas.1817066116
Adriaense, J.E.C., Martin, J.S., Schiestl, M., Lamm, C. & Bugnyar, T. 2019. Reply to Vonk: Disentangling emotional contagion from its underlying causes. Proceedings of the National Academy of Sciences of the United States of America, 116, 18169-18170. 10.1073/pnas.1910556116
Beinhauer, I., Bugnyar, T. & Auersperg, A.M.I. 2019. Prospective but not retrospective tool selection in the Goffin’s cockatoo (Cacatua goffiniana). Behaviour, 156, 633-656. 10.1163/1568539X-00003515
Boucherie, P.H., Loretto, M.-C., Massen, J.J.M. & Bugnyar, T. 2019. What constitutes ‘social complexity’ and ‘social intelligence’ in birds? Lessons from ravens. Behavioral Ecology and Sociobiology, 73 (1), 12. 10.1007/s00265-018-2607-2
Bugnyar, T. 2019. Tool use: New Caledonian crows engage in mental planning. Current Biology, 29, R200-202. 10.1016/j.cub.2019.01.059
Keupp, S., Titchener, R., Bugnyar, T., Mussweiler, T. & Fischer, J. 2019. Competition is crucial for social comparison processes in longtailed macaques. Biology Letters, 15, 20180784. 10.1098/rsbl.2018.0784
Laumer, I.B., Auersperg, A.M.I., Bugnyar, T. & Call J. 2019. Orangutans (Pongo abelii) make flexible decisions relative to reward quality and tool functionality in a multi-dimensional tool-use task. Plos One, 14, 0211031. 10.1371/journal.pone.0211031
Martin, J.S., Massen, J.J.M., Slipogor, V., Bugnyar, T., Jaeggi, A.V. & Koski, S.E. 2018. The EGA+GNM framework: an integrative approach to modelling behavioural syndromes. Methods in Ecology and Evolution, 10, 245-257. https://doi.org/10.1111/2041-210X.13100
Reber, S.A., Šlipogor, V., Oh, J., Ravignani, A., Hoeschele, M., Bugnyar, T. & Fitch, T.F. (2019) Common marmosets (Callithrix jacchus) are sensitive to simple dependencies at variable distances in an artificial grammar. Evolution and Human Behaviour, 40, 214-221. doi: 10.1016/j.evolhumbehav.2018.11.006
Uhl, F., Ringler, M., Miller, R., Deventer, S., Bugnyar, T. & Schwab, C. 2019. Counting crows: Population structure and group size variation in an urban population of crows. Behavioral Ecology, 30, 57-67. 10.1093/beheco/ary157
2018
Boeckle, M., Szipl, G. & Bugnyar, T. 2018. Raven food calls indicate sender’s age and sex. Frontiers in Zoology, 15, 5. 10.1186/s12983-018-0255-z
Horn, L., Hungerlaender, N.A., Windhager, S., Bugnyar, T. & Massen, J.J.M. 2018. Social status and prenatal testosterone exposure assessed via second-to-fourth digit ratio affect 6-9-year-old children’s prosocial choices. Scientific Reports, 8, 9198. doi: 10.1038/s41598-018-27468-0
Laumer, I.B., Call, J., Bugnyar, T. & Auersperg, A.M.I. 2018. Spontaneous innovation of hook-bending and unbending in orangutans (Pongo abelii). Scientific Reports, 8, 16518. 10.1038/s41598-018-34607-0
Marchand, P., Loretto, M.-C., Henry, P.-Y., Duriez, O., Jiguet, F., Bugnyar, T. & Itty, C. 2018. Relocations and one-time disturbance fail to sustainably disperse non-breeding common ravens Corvus corax due to homing behaviour and extensive home ranges. European Journal of Wildlife Research, 64 (5), 57. 10.1007/s10344-018-1217-7
Nacarova, J., Vesely, P. & Bugnyar, T. 2018. Ravens adjust their antipredator responses to con- and hetero-specific alarms to the perceived threat. Ethology, 124, 609-616. https://doi.org/10.1111/eth.12764
Oh, J., Reber, S.A., Slipogor, V., Hoesele, M., Bugnyar, T. & Fitch, W.T. 2018. A technological framework for running and analyzing animal head turning experiments. Behaviour Research Methods, 50, 1154-1165. 10.3758/s13428-017-0934-2
Sima, M.J., Matzinger, T., Bugnyar, T. & Pika, S. 2018. Reconciliation and third-party affiliation in carrion crows. Ethology, 124, 33-44. https://doi.org/10.1111/eth.12699
Szipl, G., Ringler, E. & Bugnyar, T. 2018. Attacked ravens flexibly adjust signaling behavior according to audience composition. Proceedings of the Royal Society London Series B, 285, 20180375. 10.1098/rspb.2018.0375
2017
Bila, K., Berankova, J., Vesely, P., Bugnyar, T. & Schwab, C. 2017. Responses of urban crows to con- and hetero-specific alarm calls in predator and non-predator zoo enclosures. Animal Cognition, 20, 43-51.
Bugnyar, T. 2017. Apes perform like infants in false belief task. Learning & Behaviour, 45, 325-326
Bugnyar, T. & Massen, J.J.M. 2017. Avian social relations, social cognition and cooperation. In Healy, S. & ten Cate, C.J. (eds) Avian Cognition, pp. 314-336, Cambridge, UK, Cambridge University Press.
Federspiel I.G., Garland, A., Guez, D., Bugnyar, T., Healy, S.D., Güntürkün, O. & Griffin, A.S. 2017. Adjusting foraging strategies: a comparison of rural and urban common mynahs (Acridotheres tristis). Animal Cognition, 20, 65-74.
Griffin, A.S., Tebbich, S. & Bugnyar, T. 2017. Animal cognition in a human-dominated world. Animal Cognition, 20, 7-7.
Koski, S.E., Buchanan-Smith, H.M., Ash, H., Burkart, J.M., Bugnyar, T. & Weiss, A. 2017. Common marmoset personality. Journal of Comparative Psychology, 131, 326-336.
Lambert, M., Massen, J.J.M., Seed, A., Bugnyar, T. & Slocombe, K. 2017. An ‘unkindness’ of ravens? Measuring prosocial preferences in Corvus corax. Animal Behaviour, 123, 383-393.
Laumer, I., Bugnyar, T., Reber, S. & Auersperg, A. 2017. Can hook-bending be let off the hook? Bending/unbending of pliant tools by cockatoos. Proceedings of the Royal Society B, 284, 20171026
Loretto, M.-C., Schuster, R., Itty, C., Marchand, P., Genero, F. & Bugnyar, T. 2017. Fission-fusion dynamics over large distances in raven non-breeders. Scientific Reports, 7, 380.
Massen, J.J.M., Bauer, L., Spurny, B., Bugnyar, T., & Kret, M.E. 2017. Sharing of science is most likely among male scientists. Scientific Reports, 7, 12927.
Munteanu, A.M., Stocker, M., Stöwe, M. & Bugnyar, T. 2017. Behavioural and hormonal stress responses to social separation in ravens, Corvus corax. Ethology, 122, 1-13.
Müller, J.J.A., Massen, J.J.M., Bugnyar, T. & Osvath, M. 2017 Ravens remember the nature of a single reciprocal interaction sequence over 2 days and even after a month. Animal Behaviour, 128, 69-78.
O’Hara, M., Mioduszewska, B., von Bayern, A., Auersperg, A., Bugnyar, T., Wilkinson, A., Huber, L. & Gajdon, G.K. 2017. The temporal dependence of exploration on neotic style in birds. Scientific Reports, 7, 4742.
Schwing, R., Weber, S. & Bugnyar, T. 2017. Kea (Nestor notabilis) decide early when to wait in food exchange task. Journal of Comparative Psychology, 131, 269-276.
Szabo, B., Bugnyar, T., Auersperg A.M.I. 2017.Within-group relationships and lack of social enhancement during object manipulation in captive Goffin’s cockatoos (Cacatua goffiniana). Learning & Behavior, 45, 7-19.
Szipl, G., Ringler, E., Spreafico, M. & Bugnyar, T. 2017. Calls during agonistic interactions vary with arousal and raise audience attention in ravens. Frontiers in Zoology, 14, 57.
2016
Asakawa-Haas, K., Schiestl, M., Bugnyar, T. & Massen, J.J.M. 2016. Partner choice in raven (Corvus corax) cooperation. Plos One 11, e0156962.
Bugnyar, T., Reber, S.A. & Buckner, C. 2016. Ravens attribute visual access to unseen competitors. Nature Communications, 7, 10506.
Deventer, S., Uhl, F., Bugnyar, T., Miller, R., Fitch, W.T.S., Schiestl., M., Ringler, M. & Schwab, C. 2016. Behavioural type affects space use in a wild population of crows (Corvus corone). Ethology, 122, 881-891.
Güntürkün, O. & Bugnyar, T. 2016. Cognition without cortex. Trends in Cognitive Sciences, 20, 291-303.
Horn, L., Scheer, C., Bugnyar. T. & Massen, J.J.M. 2016. Proactive prosociality in a cooperatively breeding corvid, the azure-winged magpie (Cyanopica cyana). Biology Letters, 12, 20160649.
Kulhaci, I.G., Rubenstein, D.I., Bugnyar, T., Hoppitt, W., Mikus, N. & Schwab, C.2016.Social networks predict selective observation and information spread in ravens. Royal Society Open Science, 3, 160256.
Laumer, I., Bugnyar, T. & Auersperg, A.M.I. 2016. Flexible decision-making relative to reward quality and tool functionality in Goffin cockatoos (Cacatua goffiniana). Scientific reports, 6, 28380.
Loretto, M.-C., Reimann, S., Schuster, R., Graulich, D. & Bugnyar, T. 2016.Shared space-individually used: spatial behaviour of non-breeding ravens (Corvus corax) close to a permanent anthropogenic food source. Journal of Ornithology, 157, 439-450.
Loretto, M.-C., Schuster, R. & Bugnyar, T. 2016. GPS tracking of non-breeding ravens reveals importance of anthropogenic food sources during their dispersal in the Eastern Alps. Curr Zool, 62, 337–344.
Miller, R.. Laskowski, K.L., Schiestl, M., Bugnyar, T. & Schwab, C. 2016. Socially driven consistent behavioural differences during development in common ravens and carrion crows. Plos One, 11, e0148822.
Miller, R., Schwab, C. & Bugnyar, T. 2016. Explorative innovators and flexible use of social information in common ravens (Corvus corax) and carrion crows (Corvus corone). Journal of comparative Psychology, 130, 328-340.
Munteanu, A.M., Starnberger, I., Pasukonis, A., Bugnyar, T., Hödl, W. & Fitch, W.T.S. 2016. Take the long way home: detour behaviour in a neotropical frog, Allobathes femoralis. Behavioural Processes, 126, 71-75.
Reber, S.A., Szipl, G., Janisch, J., Boeckle, M. Bugnyar, T. & Fitch, W.T.S. 2016.Territorial raven pairs are sensitive to structural changes in simulated acoustical displays of conspecifics. Animal Behaviour, 116, 153-162.
Schmitt, V., Federspiel, I., Eckert, J., Keupp, S., Tschernek, L., Faraut, L., Schuster, R., Michels, C., Sennhenn-Reulen, H., Bugnyar, T., Mussweiler, T. & Fischer, J. 2016. Do monkeys compare themselves to others? Animal Cognition, 19, 417-428.
Sima, M.J., Pika, S. & Bugnyar, T. 2016. Experimental manipulation of food accessibility affects conflict management behaviour in ravens. Ethology, 122, 114-126.
Slipogor, V., Gunhold-de Oliveira, T., Tadic, Z., Massen, J.J.M. & Bugnyar, T. 2016. Consistent inter-individual differences in common marmosets (Callithrix jacchus) in boldness-shyness, stress-activity, and exploration-avoidance. American Journal of Primatology, 78, 961-973
Stocker, M., Munteanu, A., Stöwe, M., Schwab, C., Palme R. & Bugnyar, T. 2016. Loner or socializer? Adrenocortical response to individual separation depends on social integration in ravens. Hormones & Behaviour, 78, 194-199.
2015
Auersperg, A.M.I., van Horik, J.O., Bugnyar, T., Kacelnik, A., Emery, N.J. & von Bayern, A.M.I. 2015. Combinatory actions during object play in psittaciformes (Diopsittaca nobilis, Pionites melanocephala, Cacatua goffini) and corvids (Corvus corax, C. monedula, C. moneduloides). Journal of Comparative Psychology, 129, 62-71.
Gunhold, T., Range, F., Huber, L. & Bugnyar, T. 2015. Long-term fidelity of foraging techniques in common marmosets (Callithrix jacchus). American Journal of Primatology, 77, 264-270.
Kubitza, R.J., Bugnyar, T. & Schwab, C. 2015. Pair-bond characteristics and maintenance in free-flying jackdaws Corvus monedula: effects of social context and season. Journal of Avian Biology, 45, 206–215.
Massen, J.J.M., Ritter, C. & Bugnyar, T.2015. Tolerance and reward equity predict cooperation in ravens. Scientific Reports, 5, 15021.
Massen, J.J.M., Lambert, M., Schiestl, M. & Bugnyar, T. 2015.Subadult ravens generally don’t transfer valuable tokens to conspecifics when there is nothing to gain for themselves. Frontiers in Comparative Psychology, 6, 885.
Miller, R., Bugnyar, T, Pölzl, K. & Schwab. C.2015. Differences in exploration behaviour in common ravens and carrion crows during development and across social context. Behavioral Ecology and Sociobiology, 69, 1209-1220.
O’Hara, M., Auerperg, A.M.I., Bugnyar, T. & Huber, L.2015. Inference by exclusion in Goffin cockatoos (Cacatua goffini). Plos One, 10, 8.
Szipl, G., Boeckle, M., Wascher, C.A.F. & Bugnyar, T. 2015. With whom to dine? Ravens’ responses to food-associated calls depend on individual characteristics of the caller. Animal Behaviour, 99, 33–42.
2014
Auersperg, A.M.I., von Bayern, A.M.I., Weber, S., Szabadvary, A., Bugnyar, T. & Kacelnik, K. 2014. Social transmission of tool use and tool manufacture in Goffin cockatoos (Cacatua goffini). Proceedings Royal Society London Series B, 281, 20140972.
Auersperg, A.M.I., Oswald, N., Domansegg, M., Gajdon, G.K., Bugnyar, T. Unrewarded object combinations in parrots. Animal Behavior and Cognition, 1 ,470-488.
Gunhold, T., Massen, J.J.M., Schiel, N., Souto, A. & Bugnyar, T. 2014. Memory, transmission and persistence of alternative foraging techniques in wild common marmosets. Animal Behaviour, 91, 79-91.
Gunhold, T., Whiten, A. & Bugnyar, T. 2014. Video demonstrations seed alternative problem solving techniques in wild common marmosets. Biology Letters, 10, 20140439.
Hillemann, F., Bugnyar, T., Kotrschal, K. & Wascher, C.A.F. 2014. Waiting for better, not for more: corvids respond to quality in two delay maintenance tasks. Animal Behaviour, 90, 1-10.
Massen, J.J.M., Pasukonis, A., Schmidt, J. & Bugnyar, T. 2014. Ravens notice dominance reversals among conspecifics within and outside their social group. Nature Communications, 5, 3679.
Massen, J.J.M., Szipl, G. Spreafico, M. & Bugnyar, T. 2014. Ravens intervene in others’ bonding attempts. Current Biology, 24, 1-4..
Miller, R., Schiestl, M., Whiten, A., Schwab, C. & Bugnyar, T. 2014. Tolerance and social facilitation in the foraging behaviour of free-ranging crows (Corvus corone corone; C.c. cornix) Ethology, 120, 1248-1255.
Pfuhl, G., Gattermayr, M. & Bugnyar, T. 2014. Will food-handling time influence agonistic behaviour in sub-adult common ravens (Corvus corax)? Behavioural Processes, 103, 67-74.
Szipl, G. & Bugnyar, T. 2014. Craving ravens: individual ‘haa’call rates at feeding sites as cues to personality and levels of fission-fusion dynamics? Animal Behavior and Cognition, 1, 265-280.
Ujfalussy, D.J., Miklosi, A., Bugnyar, T. & Kotrschal, K.2014. Role of mental representations in quantity judgments by jackdaws (Corvus monedula). Journal of Comparative Psychology, 128, 11-20.
2013
Auersperg, A.M.I, Laumer, I.B. & Bugnyar, T. 2013. Goffin cockatoos wait for qualitative and quatitative gains but prefer ‘better’ to ‘more’. Biology Letters, 9, 20121092,
Auersperg, A.M.I., Szabo, B., von Bayern, A.M.P. & Bugnyar, T. 2013. Object permanence in the Goffin Cockatoo (Cacatua goffini). Journal of Comparative Psychology, 128, 88-98.
Bugnyar, T. 2013. Social cognition in ravens. Comparative Cognition & Behavior Reviews, 8, 1-12.
Bugnyar, T. 2013. Q & A Thomas Bugnyar. Current Biology, 23, R549-R551.
Di Lascio, F., Nyffeler, F., Bshary, R. & Bugnyar, T. 2013. Ravens (Corvus corax) are indifferent to the gains of conspecific recipients or human partners in experimental tasks. Animal Cognition, 16,35-43.
Stephan, C. & Bugnyar, T. 2013. Pigeons integrate past knowledge across sensory modalities. Animal Behaviour, 85, 605-613.
Ujfalussy, D.J, Bugnyar, T., Kotrschal, K. & Miklosi, A. 2013. Ontogeny of object permanence in a non-storing corvid species, the jackdaw (Corvus monedula). Animal Cognition, 16, 405-416.
Wascher, C.A.F. & Bugnyar, T. 2013. Behavioral responses to inequity in reward distribution and working effort in crows and ravens. PloS One, 8, e56885.
2012
Albiach-Serrano, A., Bugnyar, T. & Call, J. 2012. Apes (Gorilla gorilla, Pan paniscus, P. troglodytes, Pongo pygmaeus) vs. corvids (Corvus corax, C. corone) in a support task: the effect of pattern and functionality. Journal of Comparative Psychology, 126, 355-367.
Boeckle, M. & Bugnyar, T.2012. Long-term memory for affiliates in ravens. Current Biology, 22, 801-806.
Boeckle, M., Szipl, G. & Bugnyar, T. 2012. Who wants food? Individual characteristics in raven yells. Animal Behaviour, 84, 1123–1130.
Braun, A. & Bugnyar, T.2012. Social bonds and rank acquisition in raven non-breeder aggregations. Animal Behaviour, 84, 1507-1515.
Braun, A., Walsdorff, T., Fraser, O.N. & Bugnyar, T. 2012. Socialized sub-groups in a temporary stable raven flock? Journal of Ornithology, 153, 97-104.
Bugnyar, T., Boyd, R., Bossan, B., Gächter, S., Griffiths, T., Hammerstein, P., Jensen, K., Mussweiler, T., Nagel, R. & Warneken, F. 2012. Evolutionary perspectives on social cognition. In: Evolution and the Mechanisms of Decision Making (ed. Hammerstein, P. & Stevens, J.), pp. 345-367, Ernst Strüngmann Forum Reports. MIT Press, Cambridge MA.
Dufour, V., Wascher, C., Braun, A., Miller, R. & Bugnyar, T. 2012. Corvids can decide if a future exchange is worth waiting for. Biology Letters, 8, 201-204.
Fraser, O.N. & Bugnyar, T. 2012. Reciprocity of agonistic support in ravens. Animal Behaviour, 83, 171-177.
Loretto, M.-C., Fraser, O.N. & Bugnyar, T. 2012. Ontogeny of social relations and coalition formation in common ravens (Corvus corax). International Journal of Comparative Psychology, 25, 180-194.
Schwab, C., Swoboda, R., Kotrschal, K. & Bugnyar, T.2012. Recipients affect prosocial and altruistic choices in jackdaws. PLoS One, 7, e34922.
Wascher, C.A.F., Dufour, V. & Bugnyar, T. 2012. Carrion crows cannot overcome impulsive choice in a quantitative exchange task. Frontiers in Comparative Psychology, 3(118).
2011
Bugnyar, T. 2011. Knower–guesser differentiation in ravens: others’ viewpoints matter. Proceedings Royal Society London Series B, 278, 634-640.
Fraser, O.N. & Bugnyar, T. 2011. Ravens reconcile after aggressive conflicts with valuable partners. PLoS One, 6, e18118.
Kenward, B., Schloegl, C., Weir, A.A.S., Rutz, C., Bugnyar, T. & Kacelnik, A. 2011. On the evolutionary and ontogenetic origins of tool-oriented behaviour in New Caledonian crows (Corvus moneduloides). Biological Journal of the Linnean Society, 102, 870-877.
Pika, S. & Bugnyar, T.2011. The use of referential gestures in ravens (Corvus corax) in the wild. Nature Communications, 2, 560.
Schmidt, J., Scheid, C., Kotrschal, K., Bugnyar, T. & Schloegl, C. 2011. Gaze direction – a cue for hidden food in rooks (Corvus frugilegus)? Behavioural Processes, 88, 88-93.
2010
Fitch, W.T.S., Huber, L. &Bugnyar, T.2010. Social cognition and the evolution of language: constructing cognitive phylogenies. Neuron, 65, 795-814.
Fraser, O.N. & Bugnyar, T. 2010. The quality of social relationships in ravens. Animal Behaviour, 79, 927-933.
Fraser, O.N. & Bugnyar, T. 2010. Do ravens show consolation? Responses to distressed other. PLoS One, 5, e10605.
Loretto, M.-C., Schloegl, C. & Bugnyar, T. 2010. Norther bald ibises follow others’ gaze into space but not behind barriers. Biology Letters, 6, 14-17.
Wilkinson, A., Bugnyar, T.& Huber, L. 2010. Gaze-following in the red-footed tortoise (Geochelone carbonaria). Animal Cognition, 13, 765-769.
2009
Range, F., Horn, L., Bugnyar, T., Gajdon, G. & Huber, L. 2009. Observing behaviour in keas, dogs, and human children. Animal Cognition, 12, 181-192.
Schloegl, C., Bugnyar, T. & U. Aust. 2009. Exclusion performances in non-human animals: from pigeons to chimpanzees and back again. In: Rational Animals, Irrational Humans (ed. Blaisdell, A., Huber, L., Watanabe, S., Young, A. & Yamazaki, Y.), pp. 217-234, Tokyo: Keio University Press.
Schloegl, C., Dierks, A., Gaydon, G.K., Huber, L., Kotrschal, K. & Bugnyar, T. 2009. What you see is what you get? Inference by exclusion in ravens (Corvus corax) and keas (Nestor notabilis). PLoS One, 4, e6368.
2008
Bugnyar, T. 2008. Animal cognition: Rooks solve cooperation task. Current Biology, 18, 530-532.
Bugnyar, T. 2008. Book review Reznikova, Z. ‘Animal Intelligence. From individual to social cognition’. Myrmecological News, 11, 42.
Range, F., Bugnyar, T. & Kotrschal, K. 2008. Simple task discriminations in ravens. Acta Ethologica, 11, 34-41.
Scheid, C. & Bugnyar, T.2008. Short-term observational spatial memory in jackdaws and ravens. Animal Cognition, 11, 691-698.
Schloegl, C., Kotrschal, K. & Bugnyar, T. 2008. Do common ravens (Corvus corax) rely on gaze cues to detect hidden food? Animal Cognition, 11, 231-241.
Schloegl, C., Kotrschal, K. & Bugnyar, T.2008.Modifying the object-choice task: is the way you look important for ravens? Behavioural Processes, 77, 61-65.
Schloegl, C., Schmidt, J., Scheid, C., Kotrschal, K. & Bugnyar, T. 2008. Gaze following in non-human animals: the corvid example. In: Animal Behaviour: New Research (ed. Columbus, F.), pp. 73-92, New York: Nova Science Publishers.
Schwab, C., Bugnyar, T., Schlögl, C. & Kotrschal, K. 2008. Enhanced social learning between siblings in common ravens (Corvus corax). Animal Behaviour, 75, 501-508.
Schwab, C., Bugnyar, T.& Kotrschal, K. 2008. Learning from non-affiliated individuals as a relevant source of information in jackdaws (Corvus monedula). Behavioural Processes, 79, 148-155.
Stöwe, M., Bugnyar, T., Schloegl, C., Heinrich, B., Kotrschal, K. & Möstl, E. 2008. Corticosterone excretion patterns and affiliative behaviour over development in ravens. Hormones & Behaviour, 53, 208-216.
2007
Bugnyar, T. 2007. An integrative approach to the study of ToM-like abilities in ravens. Japanese Journal of Animal Psychology, 57, 15-27.
Bugnyar, T., Schwab, C., Schlögl, C., Kotrschal, K. & Heinrich, B. 2007. Ravens judge competitors through experience with play caching. Current Biology, 17, 1804-1808.
Bugnyar, T., Stöwe, M. & Heinrich, B. 2007. The ontogeny of caching in ravens, Corvus corax. Animal Behaviour, 74, 757-767.
Heinrich, B. & Bugnyar, T. 2007. Raven intelligence. Scientific American, 296, 46-53.
Scheid, C., Range, F. & Bugnyar, T. 2007. When, what, and whom to watch? Quantitive measures of attention to conspecifics in ravens (Corvus corax) and jackdaws (Corvus monedula). Journal of Comparative Psychology, 121, 380-386.
Schlögl, C., Kotrschal, K. & Bugnyar, T. 2007. Gaze following in common ravens (Corvus corax): ontogeny and habituation. Animal Behaviour, 74, 769-778.
2006
Bugnyar, T. & Heinrich, B. 2006. Pilfering ravens, Corvus corax, adjust their behaviour to social context and identity of competitors. Animal Cognition 9, 369-376.
Range, F., Bugnyar, T., Schlögl, C. & Kotrschal, K. 2006. Individual learning ability and coping styles in ravens. Behavioural Processes, 73, 100-106.
Stöwe, M., Bugnyar, T., Heinrich, B. & Kotrschal, K. 2006. Effects of group size on exploration in ravens (Corvus corax). Ethology 112, 1079-1088.
Stöwe, M., Bugnyar, T., Loretto, M.-C., Schlögl, C., Range, F. & Kotrschal, K. 2006. Novel object exploration in ravens (Corvus corax):effects of social relationships. Behavioural Processes 73, 68-75.
2005
Bugnyar, T. & Heinrich, B. 2005. Food-storing ravens differentiate between knowlegeable and ignorant competitors. Proceedings Royal Society London Series B 272, 1641-1646.
Heinrich, B. & Bugnyar, T. 2005. Testing problem solving in ravens: String-pulling to reach food. Ethology 111, 962-976.
2004
Bugnyar, T., Stöwe, M. & Heinrich, B. 2004. Ravens, Corvus corax, follow gaze direction of humans around obstacles. Proceedings Royal Society London Series B 271, 1331-1336.
Bugnyar, T. & Kotrschal K. 2004. Leading a conspecific away from food in ravens, Corvus corax? Animal Cognition 7, 69-76.
2002
Bugnyar, T. & Kotrschal, K. 2002. Observational learning and the raiding of food caches in ravens, Corvus corax: Is it “tactical deception”? Animal Behaviour 64, 185-195.
Bugnyar, T. & Kotrschal K. 2002. Scrounging tactics in free-ranging ravens. Ethology 108, 993-1009.
2001
Bugnyar, T., Kijne, M. & Kotrschal, K. 2001. Food calling in ravens: are yells referential signals? Animal Behaviour 61, 949-958.
Bugnyar, T. & Kotrschal, K. 2001. Movement coordination and signalling in ravens (Corvus corax): an experimental field study. Acta Ethologica 3, 101-109.
1997
Bugnyar, T. & Huber, L. 1997. Push or pull: An experimental study on imitation in common marmosets (Callithrix jacchus). Animal Behaviour 54, 817-831.
Bugnyar.Lab 